Reminder that “struggle for existence” is a conceptual metaphor.
In the November, 2009 edition of Nature (available only to subscribers online) Daniel Todes has written an article entitled "Global Darwin: Contempt for Competition." Todes points out that although Darwin's idea of a "struggle for existence" made sense to his English peers, other biologists from other countries rejected this metaphor. Todes focuses on the alternative viewpoint embraced by many Russian biologists. In On the Origin of Species (1859), Darwin noted that "there must in every case be a struggle for existence, either one individual with another of the same species, or with the individuals of distinct species, or with the physical conditions of life. It is the doctrine of Malthus applied with manifold force to the whole animal and vegetable kingdoms." Through this quote, Darwin recognized that he was using "struggle for existence" in a metaphorical sense. Darwin had been urged to adopt this metaphor by Russel Wallace, who feared that natural selection "seemed to personify a perceptive and forward thinking selector, or god." Todes holds that Darwin's metaphor was common sense to those "who were living on a crowded island with a capitalist economy and highly individualist culture." Russian biologists lived in a very different place, however, which led them to "reject Darwin's Malthusian metaphor." Russians did not tend to explore densely populated tropical environments. Rather, they tended to investigate "a vast underpopulated continental plain . . . it was largely empty Siberian expanse in which overpopulation was rare and only the struggle of organisms against a harsh environment was dramatic." Todes points out the Russian political system also contrasted sharply with that of Darwin's England. In Russia, capitalism was only weakly developed, and the social classes stressed cooperation rather than individual struggle, one against the other. In fact, many Russian political commentators "reviled Malthus as an apologist for predatory capitalism and the soulless individualism." This context for the Russian research led to (many successful) studies in which the focus was "mutual aid" more than "struggle for existence." [more . . . ]